Stems erect, 2-8 dm long, diffusely branched, pubescent with long, spreading and also short, appressed hairs. Leaves elliptic to ovate, 2-10 cm long, 1-7 cm wide, with glands on lower surface, margins serrate to subentire, gradually narrowed to base, sessile. Heads 4-5 mm in diameter at anthesis. Burs cuneate, strongly compressed, 4-7 mm long, the ribs bearing 1-2 rows of hooked prickles 1-2 mm long, the 2 apical prickles stout, curved or straight, 3-4 mm long." (Wagner et al., 1999). |